Semple, J. C. 1984. Cytogeographic studies on North American Asters. I. Range surveys of Virgulus adnatus, V. concolor, V. georgianus, V. grandiflorus, V. novae-angliae, V. oblongifolius, V. patens and V. walteri. Amer. J. Bot. 71: 522–531.
Semple, J. C., J. G. Chmielewski & C. Xiang. 1992. Chromosome number determinations in fam. Compositae, tribe Astereae. IV. Additional reports and comments on the cytogeography and status of some species of Aster and Solidago. Rhodora 94: 48–62.
Semple, J. C., J. Zhang & C. Xiang. 1993. Chromosome number determinations in fam. Compositae, tribe Astereae. V. Eastern North American taxa. Rhodora 95: 234–253.